摘要
The rhipidoglossan radula, which is characterized by presence of a central tooth, several lateral teeth, and numerous (more than 10) long marginal teeth in each transverse row, is found in three different subclasses, that is, Vetigastropoda, Neritomorpha and “lower” Heterobranchia. Details of radula formation and its ultrastructure have not been studied in any species with a rhipidoglossan radula. For the first time, we present such data for one vetigastropod species, Puncturella noachina. The radula itself and the radula formation zone were studied using light and electron microscopy (scanning and transmission), as well as confocal laser scanning microscopy. We identify the major features of Vetigastropoda rhipidoglossan radula formation, that is: the posterior bifurcation of the radula formation zone, creating paired horns into which the zones of formation of the marginal teeth extend; the supporting structure in the radula formation zone extends ventrally to strengthen this division; the odontoblasts of the marginal teeth form a multi-layered epithelium; membranoblasts do not differ from odontoblasts in ultrastructure; in some membranoblasts and cells of the sub- and supraradular epithelium basal bodies were found in the apical regions of the cells.
摘要译文
在三种不同的亚类中发现了在每个横向行中存在中央牙齿，几个横向牙齿和许多（超过10个）的长边缘齿的refhidoglossan adrula，即vetigaStropoda，Neritomorpha和“下”异常。罗格拉米龙罗拉罗布的任何物种都没有研究罗布拉米菌的细节及其超微结构。我们首次为vetigafetropod种类，Puncturella Noachina提供此类数据。使用光和电子显微镜（扫描和透射）研究罗格拉本身和射线形成区，以及共聚焦激光扫描显微镜。我们确定了vetigastrophoda rhipidoglossan adrula形成的主要特点，即：射迹形成区的后部分叉，产生配对角，边缘齿的形成区域延伸到其中;射频形成区的支撑结构介绍以加强该划分;边缘牙齿的成牙细胞形成多层上皮;膜质细胞与超微结构中的牙卵细胞不同;在细胞的顶端区域中发现了在一些膜和寄生上皮基体的细胞中。
Elena Vortsepneva[1];David G. Herbert[2];Yuri Kantor[3]. The rhipidoglossan radula: Formation and morphology of the radula in Puncturella noachina (Linnaeus, 1771) (Fissurellidae, Vetigastropoda)[J]. Journal of Morphology, 2021,282(10): 1523-1532